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Did you know Rob Lowe almost played Derek Shepherd in Grey's Anatomy?
While the role ultimately went to Kit Harington, Rheon's talent kept him on the showrunners' minds. They ended up casting him as Ramsay Bolton in Season 3.
After being offered the role of Rachel, Cox said that she would rather play Monica.
Both Broderick and John Cusack turned down the role of Walter White before it was offered to Bryan Cranston.
She was originally considered for the role before Sarah Michelle Gellar booked the gig.
He was offered the role of McDreamy, but turned it down to star in another series, CBS's Dr. Vegas.
Retta said that when she was called in to audition, Spencer was also among the contenders for the role.
She originally auditioned for the role of Kelly before the role went to Tiffany Thiessen. Garth ended up winning her iconic role as Kelly Taylor in Beverly Hills, 90210 instead.
Casting director David Rapaport said that Willis was considered for the role, but he ultimately felt that Blake Lively, a lesser known actor at the time, fit the show's vibe more.
Chris Rock, Danny DeVito, and numerous others all turned down the role of George before Jason Alexander was cast.
He ultimately turned down the role, saying that he knew it wasn't the right fit for him.
She was offered the role of Robin, but ended up turning it down for her role in Ghost Whisperer.
The youngest Huxtable kid was originally envisioned to be a young boy. White ended up booking his iconic role of Steve Urkel instead.
Originally, Jack was supposed to be shockingly killed off in the pilot episode. When they decided to keep Jack on as a series regular, Keaton bowed out of the project.
She originally auditioned for the role of Michonne before she was ultimately cast as Sasha in the hit series.
The young actors both auditioned for each other's roles in the short-lived Boy Meets World spin-off series.
He originally auditioned for the role of Jim before the role went to John Krasinski.
Silverstone and Claire Danes were the two up-and-coming actors called into audition for the show. The role ultimately went to Danes.
She originally auditioned for the role of Peggy before she was ultimately cast in the role of Rachel Menken.
She was originally slated to be Sookie St. James, but ended up having to drop out of the project because of her contract with MadTV. They even filmed an unaired pilot episode with Borstein as Sookie.
He revealed that he auditioned for all three leading men in Full House and was ultimately turned down.