1. Evan Williams, co-founder: 12%, 56.9 million shares
2. Jack Dorsey, co-founder and executive chairman: 4.9%, 23 million shares
3. Dick Costolo, Twitter CEO: 1.6%, 7.6 million shares
Matthew Zeitlin is a business reporter for BuzzFeed News and is based in New York. Zeitlin reports on Wall Street and big banks.
Contact Matthew Zeitlin at firstname.lastname@example.org.
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